I was a PhD student at the Multisensory Integration lab. My research interests lie in the fields of multisensory integration, attention and sensory-motor sequence learning. My project focused on the neural and behavioral effects of attention and expectations on audio-visual integration. After my PhD I started working at flowkey, an innovative company that developes and provides new approaches to learn musical instruments.
Professional experience
Since 2022       Engineering for flowkey (
Academic Background
Since 2017       PhD student at 'Multisensory Integration' group, Charité Universitätsmedizin Berlin
2015 - 2016      Research assistant at Neurophysics Group, Charité Universitätsmedizin Berlin
2013 - 2015      Master of Science in Medical Neurosciences, Charité Universitätsmedizin Berlin
2010 - 2013      Bachelor of Science in Biological Sciences, University of Cyprus, Nicosia
Michail G., Toran Jenner L., Keil J. (2021) Prestimulus alpha power but not phase influences visual discrimination of long-duration visual stimuli. Eur J Neurosci.
Michail G., Senkowski D., Niedeggen M., Keil J. (2021) Memory Load Alters Perception-Related Neural Oscillations during Multisensory Integration. J Neurosci 41:1505–1515.
jmoran portrait edit2
I am a post-doctoral researcher interested generally in neuropsychology of clinical psychological disorders. My present role involves work on multisensory processing in Schizophrenia, using EEG; considering the potentially unique manner in which people with schizophrenia perceive the external world, and how better understanding this could help us to explain aspects of the symptoms and aetiology of the disorder. I have also studied PTSD and aggression using MEEG, as well as examining olfactory and endocrinological aspects of these phenomena.
I am currently conducting my own DFG-funded project (GEPRIS) looking at the way memory is processed in people with PTSD, using EEG and tACS to examine the neurophysiological correlates of contextual memory processing.


Dr. rer. nat. (University of Konstanz);
Master of Science (Psychology), University of Konstanz
B. Psych(Hons) (Macquarie University, Sydney)


Since 2017: Charité Medical School Berlin; Learning and Memory; M24 Course Neuroimaging of Affective Disorders
SS 2016 Uni Konstanz: "Diagnostic Assessment of PTSD" (Master) (2SWS)
WS 2015-2016: "Brain Development across the Lifespan and Plasticity" (Bachelor) (2SWS)
"PTSD Diagnostics" (Master)


Selected Publications:
Senkowski, D., & Moran, J. K. (2022). Early evoked brain activity underlies auditory and audiovisual speech recognition deficits in schizophrenia. NeuroImage: Clinical,
Keil, J., Senkowski, D., & Moran, J. K. (2022). Early and late evoked brain responses differentially reflect feature encoding and perception in the flash-lag illusion. NeuroImage, 246, 118787.
Moran, J. K., Keil, J., Masurovsky, A., Gutwinski, S., Montag, C., & Senkowski, D., (2021). Multisensory Processing can compensate for top-down attention deficits in Schizophrenia. Cerebral Cortex, 31(12), 5536–5548,
Gutwinski, S., Westerbarkey, E., Schouler-Ocak, M., Moran, J. K., & Schreiter, S. (2021) Housing Satisfaction of Psychiatric Patients in Different Forms of Housing-A Cross-Sectional Study in Berlin, Germany. Frontiers in Psychiatry, 12.
Moran, J. K., Bretz, J., Winkler, J., Gutwinski, S., Brandl, E. J., & Schouler-Ocak, M. (2021) The Differential Impact of Lockdown Measures Upon Migrant and Female Psychiatric Patients – A Cross-Sectional Survey in a Psychiatric Hospital in Berlin, Germany. Frontiers in Psychiatry, 12,
Winkler, J. G., Masah, D. J., Moran, J. K., Bretz, J., Tsagkas, I., Goldschmidt, T., & Schouler-Ocak, M. (2021). Psychische Belastung während der COVID-19-Pandemie: Konsequenzen für psychiatrisch Erkrankte und therapeutische Implikationen. Der Nervenarzt, 1-9.
Speer, L., Schuler, M., Keil, J., Moran, J. K., Pantazidis, P., Amelung, T., ... & Senkowski, D. (2020). Sexual preference for prepubescent children is associated with enhanced processing of child faces in juveniles. European Child & Adolescent Psychiatry, 1-14.
Moran, J. K., Michail, G., Heinz, A., Keil, J., & Senkowski, D. (2019). Long-range temporal correlations in resting state beta oscillations are reduced in schizophrenia. Frontiers in Psychiatry, 10, 517.
Orang, T, Ayoughi, S., Moran, J. K., Ghaffari, H., Mostafavi, S., Rasoulian, M., & Elbert, T. (2018). The efficacy of Narrative Exposure Thereapy (NET) in a sample of Iranian women exposed to ongoing intimate partner violence (IPV); A randomized controlled trial. Journal of Clinical Psychology and Psychotherapy, 25(6), 827-841.
Elbert, T., Schauer, M., & Moran, J. K. (2018). Two pedals drive the bi-cycle of violence: reactive and appetitive aggression. Current Opinion in Psychology.
Moran, J. K., Crombach, A., Elbert, T., Nandi, C., Bambonyé, M., Wienbruch, C., Lommen, U., Weierstall, R. (2017). The individual contribution of DSM 5 symptom clusters of PTSD, life events, and childhood adversity to frontal oscillatory brain asymmetry in a large sample of active combatants. Biological Psychology, 129, 305-313.
Haer, R., Hermenau, K., Elbert, T., Moran, J. K., & Hecker, T. (2017). The role of intrinsic and extrinsic rewards in committing violence during combat: A cross-sectional study with former combatants in the DR Congo. Aggressive Behavior.
Schalinski, I., Moran, J. K., Elbert, T., Reindl, V., & Wienbruch, C. (2017). Oscillatory Magnetic Brain Activity is Related to Dissociative Symptoms and Childhood Adversities–A Study in Women with Multiple Trauma. Journal of Affective Disorders. 218, 428-436.
Elbert, T., Moran, J. K., & Schauer, M. (2017). Lust an Gewalt: appetitive Aggression als Teil der menschlichen Natur. e-Neuroforum, 23(2), 96-104.
Elbert, T., Moran, J. K., & Schauer, M. (2016). Appetitive Aggression, in B. J. Bushman (Ed.) "Agression and Violence: A Social Psychological Perspective (Frontiers of Social Psychology)", Routledge: New York.
Moran, J. K., Dietrich, D. R., Elbert, T., Pause, B. M., Kübler, L., & Weierstall, R. (2015). The Scent of Blood: A Driver of Human Behavior?. PloS one. 10(9), e137777.
Moran, J., Weierstall, R., & Elbert, T. (2014). Differences in brain circuitry for appetitive and reactive aggression as revealed by realistic auditory scripts. Frontiers in Behavioral Neuroscience, 8. 425.
Schalinski, I., Moran, J., Schauer, M., & Elbert, T. (2014). Rapid emotional processing in relation to trauma-related symptoms as revealed by magnetic source imaging. BMC Psychiatry, 14(1), 193.



Dr. James Moran,
AG Multisensorische Integration
Psychiatrische Universitätsklinik 
der Charité im St. Hedwig Krankenhaus
james-kenneth.moran at


daniel imrf

Academic background

2020:   Approbation in Cognitive Behavioral Therapy (CBT), Berliner Fortbildungsakademie
2016 - :      
Supervision of Neuropsychological Assessments, Charité Berlin
2015 - :   Head of the Computational Neuroimaging Division, Charité Berlin
2012 - :   Professor of Clinical Neuropsychology, Charité Berlin
2011 - :   Head of the Research Group Multisensory Integration, Charité Berlin
2010:   Habilitation and venia legendi in Psychology, University of Hamburg
2006-2011:   Head of the Research Group Elektroencephalography, University Medical Center Hamburg-Eppendorf
2005-2006:   Adjunct Professorship, Department of Psychology, City College of New York, USA

Postdoctoral fellow, Nathan S. Kline Institute für Psychiatric Research, Orangeburg, NY

2004:   Ph.D., Max Planck Institute for Human Cognitive and Brain Sciences, Leipzig
2002-2003:    Visiting researcher, Center for Cognitive Neuroscience, Duke University, North Carolina
2000:   Diploma in Psychology, Free University Berlin



Principle Investigator

Since 2023:   "Working memory dysfunction in schizophrenia: an investigation of multivariate activation pattern and neural synchrony", Co-Investigators: Paul Sauseng (LMU Munich) and Radoslaw Cichy (FU Berlin); German Research Foundation
Since 2022:   "Examining the neural network dynamics underlying the temporal window of crossmodal perception", Co-Investigator: Tim Rohe (FAU Erlangen-Nurnberg); German Research Foundation
2016 -2020:     "Role of aberrant neural oscillations for multisensory processing deficits in schizophrenia"; German Research Foundation
2014-2017:   "Neural signatures of intersensory selective attention: investigating the role of neural synchrony and functional network properties", Co-Investigator: Julian Keil (Kiel University); German Research Foundation
2014-2016:    "Analysis and Visualization of Neural Oscillations in Electrocorticographic Singals", Co-PI: Thomas Thesen (NYU); National Institute of Health and Federal Ministry of Education and Research
2011-2016:     "The multisensory mind: from neural mechanisms to cognition", ERC Starting Grant, European Research Council.
2008-2013:   "Multisensory processing of pain: An investigation of neural binding mechanisms in electrophysiological and hemodynamic brain responses", Co-Investigators: Michael Hauck, Andreas K. Engel (UKE Hamburg), German Research Foundation
2010:   "Examining speech processing in bilateral cochlear implant users", Young Investigator Research Fund of the University Medical Center Hamburg-Eppendorf



Since 2022:   "Neural signatures of contextual memory in post-traumatic stress disorder", PI: James Moran (Charite Berlin), German Research Foundation
2014-2016:     "Predicting the multisensory future: Role of neuronal connectivity for integrative information processing across modalities", PI: Julian Keil (University Kiel), German Research Foundation
2011-2012:   "Multisensory form processing in extrastriate visual cortex", PI: John J. Foxe (University of Rochester), RO1, National Science Foundation
2004-2008:   "The neurophysiology of multisensory integration", PI: John J. Foxe, RO1 National Institute of Mental Health
2004-2007:   "Electrophysiological correlates of cognitive processes: ERP and fMRI studies of the early pre-attentive mechanisms by which the auditory system organizes auditory information", Co-Applicant: John J. Foxe, RO1, National Institute of Neurological Disorders and Stroke


2017:   Replication Study Award of the Berlin Institute of Health, Multiensory Research article "Individual alpha frequency relates to the sound-induced flash illusion" by Keil and Senkowski
2010:              Best Paper of the Year, Spanish Society for Experimental Psychology, TICS article: "The multifaceted interplay between attention and multisensory integration" by Talsma, Senkowski, Soto-Faraco, Woldorff 
2006:   NARSAD Young Investigator Award, Project: The Cocktail Party Phenomenon in Schizophrenia. Brain & Behavior Research Foundation


Since 2011:    Department of Psychiatry and Psychotherapy, Charité - University Medicine Berlin  
  • Lectures: Neural Plasticity; Cognitive Neuroscience
  • Seminar: Learning and Memory; Further lectures
  • Practical Course: Physical Exercise; Neuropsychological Assessments
2012-2015:    Department of Psychology, Free University of Berlin   
  • Seminar: Neurocognitive Psychology; Experimental Psychology: Attention and Perception
Practical Course: Empirical Practicum



  Department of Biological Psychology and Neuropsychology, University Hamburg   Lectures: Introduction to Cognitive Neuroscience; Methods of Cognitive Neuroscience



  Department of Neurophysiology and Pathophysiology, University Medical Center Hamburg-Eppendorf  
  • Seminar: Neurophysiology; Vegetative Physiology; Learning; Hearing Impairments and Deafness
  • Practical Course: Electroencephalography; Vertigo and Deafness; Sensory Function of Skin and Pain;Nerves and Muscles; EMG and Reflexes
  • Review Course: Visual System
2005-2006:   Department of Psychology, City College of New York, USA   Seminar: Advanced Experimental Design; The Synaptic Self



N = 60 peer review articles (20 first author, 25 senior author)

Google Scholar (11/23): > 5000 citations, h-index = 34


Representative articles

Multisensory integration
Michail G, Senkowski D, Holtkamp M, Wachter B, Keil J. 2022. Early beta oscillations in multisensory association areas underlie crossmodal performance enhancement. Neuroimage 257:119307 (PDF)
Michail G, Senkowski D, Niedeggen M, Keil J (2021). Memory load alters perception-related oscillations during multisensory integration. Journal of Neuroscience. 41: 1505-1515 (Abstract)
  • Keil J and Senkowski D. 2018. Neural oscillations orchestrate multisensory integration. The Neuroscientist 24: 609-626. (Abstract)
Balz J, Keil J, Roa Romero Y, Mekle R, Schubert F, Aydin S, Ittermann B, Gallinat J, Senkowski D. 2016. GABA concentration in superior temporal sulcus predicts gamma power and perception in the sound-induced flash illusion. Neuroimage 125: 724-730. (Abstract)
Schneider TR, Lorenz S, Senkowski D, Engel AK. 2011. Gamma-band activity as a signature for cross-modal priming of auditory object recognition by active haptic exploration. Journal of Neuroscience. 2011. 31:2502-2510. (Abstract)
Talsma D, Senkowski D, Soto-Faraco S, Woldorff MG. 2010. The multifaceted interplay between attention and multisensory integration. Trends in Cognitive Sciences 14: 400-410. (Abstract)
Senkowski D, Schneider TR, Foxe JJ, Engel AK. 2008. Crossmodal binding by neural coherence: Implications for multisensory processing. Trends in Neurosciences 31: 401-409. (Abstract)
Senkowski D, Molholm S, Gomez-Ramirez M, Foxe JJ. 2006. Oscillatory beta activity predicts response speed during a multisensory audiovisual reaction time task: a high-density electrical mapping study. Cerebral Cortex. 16:1556-1565. (Abstract)
Senkowski D and Moran JK. 2022. Early evoked brain activity underlies auditory and audiovisual speech recognition deficits in schizophrenia. Neuroimage:Clin. 33, 102909 (PDF)
Moran JK, Keil J, Masurovsky A, Gutwinski S, Montag C, Senkowski D. 2021. Multisensory processing can compensate for top-down attention deficits in schizophrenia. Cerebral Cortex 3:5536-5548 (Abstract)
Wooldridge J, Kaiser M, Roa Romero Y, Riecke L, Keil J, Senkowski D. 2020. Low-frequency oscillations reflect aberrant tone restauration during the auditory continuity illusion in schizophrenia. Scientific Reports 10:11872 (PDF)
Roa Romero Y, Keil J, Balz J, Gallinat J, Senkowski D. 2016. Reduced frontal theta oscillations indicate altered crossmodal prediction error processing in schizophrenia. Journal of Neurophysiology 116: 1396-1407 (Abstract)
Senkowski D and Gallinat J. 2015. Dysfunctional prefrontal gamma-band oscillations reflect working memory and other cognitive deficits in schizophrenia. Biological Psychiatry 77: 1010-1019. (Abstract)
Gallinat J, Winterer G, Herrmann CS, Senkowski D. 2004. Reduced oscillatory gamma-band responses in unmedicated schizophrenic patients indicate impaired frontal network processing. Clinical Neurophysiology. 115:1863-1874 (Abstract)
Senkowski D and Heinz A. 2016. Chronic pain and distorted body image: implications for multisensory feedback interventions. Neuroscience & Biobehavioral Reviews 69: 252-259. (Abstract)
Senkowski D, Hofle M, Engel AK. 2014. Crossmodal shaping of pain: a multisensory approach to nociception. Trends in Cognitive Sciences 18: 319-327. (Abstract)
Pomper U, Hofle M, Hauck M, Kathmann N, Engel AK, Senkowski D. 2013. Crossmodal bias of visual input on pain perception and pain-induced beta activity. Neuroimage. 66:469-478. (Abstract)
Hofle M, Pomper U, Hauck M, Engel AK, Senkowski D. 2013. Spectral signatures of viewing a needle approaching one's body when anticipating pain. Eur J Neurosci. 38:3089-3098. (Abstract)
Hofle M, Hauck M, Engel AK, Senkowski D. 2012. Viewing a needle pricking a hand that you perceive as yours enhances unpleasantness of pain. Pain 153: 1074-1081. (PDF)(Press release)(Scientific American podcast)
Senkowski D, Kautz J, Hauck M, Zimmermann R, Engel AK. 2011. Emotional facial expression modulate early pain-induced beta and gamma oscillations in sensorimotor cortex. Journal of Neuroscience 31: 14542-14550. (Abstract)
Other topics
  • Senkowski D, Ziegler T, Singh M, Heinz A, He J, Silk T, Lorenz RC. 2023. Assessing inhibitory control deficits in adult ADHD: A systematic review and meta-analysis of the stop-signal task. Neuropsychol Rev. Online ahead of print (PDF)
  • Keil J, Senkowski D, Moran JK. 2022. Early and late evoked brain responses differentially reflect feature encoding and perception of the flash-lag illusion. Neuroimage. 246:118787 (PDF)
  • Speer L, Schuler M, Keil J, Moran JK, Pantazidis P, Amelung T, Florack J, Beier KM, Senkowski D. 2022. Sexual preference for prepubescent children is associated with enhanced processing of child faces in juveniles. Eur Child Adolesc Psychiatry. (Abstract)
  • Senkowski D, Pomper U, Fitzner I, Engel AK, Kral A. 2014. Beta-band activity in auditory pathways reflects speech localization and recognition in bilateral cochlear implant users. Hum Brain Mapp. 35:3107-3121. (PDF)
Senkowski D, Röttger S, Grimm S, Foxe JJ, Herrmann CS. 2005. Kanizsa subjective figures capture visual spatial attention: evidence from electrophysiological and behavioral data. Neuropsychologia. 43:872-886. (PDF)
Senkowski D, Linden M, Zubrägel D, Bär T, Gallinat J. 2003. Evidence for disturbed cortical signal processing and altered serotonergic neurotransmission in generalized anxiety disorder. Biol Psychiatry. 53:304-314 (PDF)

Books and book chapter

Keil J and Senkowski D. 2019. Neural network dynamics and audiovisual integration. In: Springer Handbook of Auditory Research on Multisensory Processes. Editor: Wallace M.
Senkowski D and Engel AK. 2012. Oscillatory activity and multisensory processing. In: The New Handbook of Multisensory Processes. MIT Press. Editor: Stein BE. MIT Press. p. 193-205.
Talsma D, Senkowski D, Soto-Faraco S, Woldorff MG. 2012. Influences of top-down attention on multisensory processing. In: The New Handbook of Multisensory Processes. MIT Press. Editor: Stein BE, p. 371-381.
Engel AK, Senkowski D, Schneider TR. 2012. Multisensory integration through neural coherence. Frontiers in the Neural Bases of Multisensory Processing. Editors: Murray MM and Wallace MT. CRC Press, p. 115-130.
Senkowski D. 2004. Neuronal correlates of selective attention: An investigation of electrophysiologial brain responses in the EEG and MEG. MPI Series for Human Cognitive and Brain Sciences; 42, Leipzig: Max Planck Institute for Human Cognitive and Brain Sciences.

Editorial work

2013- :       Review Editor, Frontiers in Neuropsychiatric Imaging and Stimulation
2010- :   Scientific Review Associate, European Journal of Neuroscience
2009:       Guest Editor, Experimental Brain Research, Special issue on Crossmodal Processing Volume 198, Numbers 2-3 / September 2009
2008- :   Review Editor, Frontiers in Integrative Neuroscience


Ad hoc reviewer

American Journal of Psychiatry; Biological Psychiatry; Biological Psychiatry Neuroimaging; Biological Psychology; BMC Neuroscience; Brain; Brain and Cognition; Brain Research; Brain Topography; Cerebral Cortex; Clinical Neurophysiology; Clinical Psychology Reviews; Cortex; Current Biology; European Journal of Neuroscience; eLife; European Journal of Pain; Experimental Brain Research; Frontiers in Aging Neuroscience; Frontiers in Human Neuroscience; Frontiers in Integrative Neuroscience; Frontiers in Neuroscience; Frontiers in Psychology; Human Brain Mapping; Human Psychopharmacology; International Journal of Psychophysiology; Journal of Cognitive Neuroscience; Journal of Neurophysiology; Journal of Neuroscience; Multisensory Research; Nature Communications; Nature Human Behaviour; Neuroimage; Neuroimage Clinical; Neuropsychologia; Neuropsychology Review; Neuroreport; Neuroscience; Neuroscience Biobehavioral Reviews; Neuroscience Letters; Plos One; Proceedings of the National Academy of Sciences; Psychonomic Bulletin & Review; Psychiatric Research; Psychonomic Bulletin & Review; Psychophysiology; Science Advances; Scientific Reports; Schizophrenia; Trends in Cognitive Sciences
Additional reviews have been provided for various national and international funding organisations (not listed here for confidentiality reasons).



Now affiliated with Science of Intelligence


Academic Background:

10/2007 - 02/2014: Diploma in Psychology, University of Konstanz

01/2013 - 06/2014: Research assistance at the Neuroelectromagnetic Oscillations Lab, University of Konstanz

08/2012 - 10/2012: Internship at the MEG Lab, Center for Mind/Brain Sciences, University of Trento, Italy

05/2011 - 07/2011: Internship at the Critical Hermeneutics Lab, York University, Toronto, Canada

09/2010 - 04/2011: Study Exchange at York University, Toronto, Canada

11/2008 - 06/2010: Research assistance at the Ongoing-Brain-Oscillations-and-Behavior Lab, University of Konstanz



Peer-reviewed articles:

Kaiser, M., Senkowski, D., Busch, N. A., Balz, J., & Keil, J. (2019). Single trial prestimulus oscillations predict perception of the sound-induced flash illusion. Scientific Reports, 9(1), 5983.

Kaiser, M., Senkowski, D., Roa Romero, Y., Riecke, L., & Keil, J. (2018). Reduced low-frequency power and phase locking reflect restoration in the auditory continuity illusion. European Journal of Neuroscience.


Conference Papers and Presentations:

Kaiser, M., Senkowski, D., Keil, J. (2016, November). Oscillatory EEG activity in the ventriloquist illusion. Poster abstract presented at the 46th annual meeting of the Society for Neuroscience, San Diego, California

Kaiser, M., Roa Romero, Y., Riecke, L., Senkowski, D., Keil, J. (2016, May). Filling-in: Interaction effect on delta brain oscillations in the auditory continuity illusion. Poster abstract presented at the 42nd Conference Psychologie & Gehirn, Berlin, Germany

Kaiser, M., Balz, J., Senkowski, D., Keil, J. (2015, June). The Role of prestimulus oscillatory power and functional connectivity in the sound-induced flash illusion. Poster presented at the 16th International Multisensory Research Forum, Pisa, Italy

Kaiser, M., Popov, T. , & Dalal, SS. (2014, August). The electroretinogram exhibits eyes-closed alpha oscillations that couple with visual cortex. Poster presented at the 19th International Conference on Biomagnetism, Halifax, Canada


Book chapters:

Kaiser, M., & Cromby, J. (2014). Neuroscience. In T. Teo (Ed.), Encyclopedia of Critical Psychology (pp. 1243–1248). New York, NY: Springer



Dipl. Psych. Mathis Kaiser
Multisensory Integration Research Group
Department of Psychiatry and Psychotherapy
St. Hedwig Hospital
Charité - Universitätsmedizin Berlin
Große Hamburger Str. 5-11
10115 Berlin


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